This post describes a position that doesn't get much details written about it out there. The second wheel is probably the most important position in the paceline, more so even than the leader. The Second Wheel calls the shots, dictates the pace, and determines the direction that the lead rider should pull off.
If the wind is coming from the right, Second Wheel should tell the Leader to pull off to the right, into the wind. All riders should be lined up at an angle to the left of the rider ahead of them, overlapping wheels. If someone were to pull of to the left, this could cause a crash. If the wind is coming from the left, then the Second Wheel rider tells the Leader to pull of to the left, into the wind.
If the leader accelerates too much on their turn, Second Wheel needs to communicate to them to keep it steady. If the pace is getting strung out the Second Wheel needs to communicate to bring the speed down. There is no "one" person who is Second Wheel, like there is no "one" person who is the leader. This position is constantly changing as the paceline rotates. So it is important that everyone on the ride is aware of the responsibilities of the Second Wheel position.
The Second Wheel rider should also communicate when "The Whip" has occurred. In this situation, the Second Wheel rider is actually the one who has just pulled off. They will be the first to see that there are no riders continuing to pull through. They need to communicate to the Leader that they will continue the paceline starting anew. This will keep the speed high and steady and eliminate surges that can cause crashes.
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